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The optimum point at which your breathing and metabolism are fully geared for burning body fat as the primary way of fuelling your body. As a general rule, you are in your THR when you do cardio and you are not breathing too quickly or heavily and when you are not able to speak as normal (but not dying!). If you can talk but need to gasp for air every few sentences, then you are in your THR. You can of course use a chart and monitoring devices to be accurate.
